Oliver Smithies grave (2017)

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Oliver Smithies

Oliver Smithies (23 June 1925 – 10 January 2017) was a British-American geneticist and physical biochemist. He is known for introducing starch as a medium for gel electrophoresis in 1955,^[4] and for the discovery, simultaneously with Mario Capecchi and Martin Evans, of the technique of homologous recombination of transgenic DNA with genomic DNA, a much more reliable method of altering animal genomes than previously used, and the technique behind gene targeting and knockout mice.^[5]^[6]^[7] He received the Nobel Prize in Physiology or Medicine in 2007 for his genetics work.^[8]^[9]

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